My Colt New Service .45 ACP has exactly the same chamber dimensions as my .455, except for the difference in cylinder length where the .45 ACP is faced off at the rear to provide clip-clearance. If .45 ACP rounds are dropped into the chambers without the clip, the case heads are flush with the rear of the cylinder and the rounds come to a stop against the forward cone of the chamber, which is identical to my .455. Contrary to popular folklore, the chambers are not bored "straight through," but have a normal conical transition from the chamber body to the .456" diameter cylinder throats. The revolver shoots quite acceptably with ordinary GI hardball.
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Early production M1917s were in-fact assembled using existing stocks of leftover .455 cylinders from the British and Canadian orders, simply facing them off to provide clip clearance, as a manufacturing expedient.
Once the excess .455 cylinders were used up, then purpose-built .45 ACP cylinders were produced having the appropriate square stop surface to headspace on the case mouth, instead of depending upon the clips, so that loose ammunition could be used in an emergency. If revolvers were recylindered during repair or refurbishment, replacement cylinders would have been those purpose-built to .45 ACP.